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 Part Five -The Patterns and Processes of Evolution 463

 22. The Mechanisms of Evolution 486

22.1 What Facts Form the Base of Our Understanding of Evolution? 487
Adaptation has two meanings 489
Population genetics provides an underpinning for Darwin’s theory 489
Most populations are genetically variable 490
Evolutionary change can be measured by allele and genotype frequencies 491
The genetic structure of a population does not change over time if certain conditions exist 492
Deviations from Hardy–Weinberg equilibrium show that evolution is occurring 493

22.2 What Are the Mechanisms of Evolutionary Change? 494
Mutations generate genetic variation 494
Gene flow may change allele frequencies 494
Genetic drift may cause large changes in small populations 494
Nonrandom mating changes genotype frequencies 495

22.3 What Evolutionary Mechanisms Result in Adaptation? 497
Natural selection produces variable results 497
Sexual selection influences reproductive success 498

22.4 How Is Genetic Variation Maintained within Populations? 501

Neutral mutations may accumulate within populations 501
Sexual recombination amplifies the number of possible genotypes 501
Frequency-dependent selection maintains genetic variation within populations 501
Environmental variation favors genetic variation 502
Much genetic variation is maintained in geographically distinct subpopulations 503

22.5 What Are the Constraints on Evolution? 503

Developmental processes constrain evolution 504
Trade-offs constrain evolution 504
Short-term and long-term evolutionary outcomes sometimes differ 504

22.6 How Have Humans Influenced Evolution? 505

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